Corylus plant named 'YORK'

Abstract

A new and distinct cultivar of Corylus plant named `York` characterized by globose plant habit and moderate vigor, green developing and fully expanded leaves during the spring and summer, resistance to eastern filbert blight caused by the fungus Anisogramma anomala (Peck) E. Muller, presence of random amplified polymorphic DNA markers 152-800 and AA12-850, expression of incompatibility alleles S₂ and S₂₁ in the styles, and DNA fingerprints at 10 of 23 microsatellite marker loci that differ from both parents OSU 479.027 and OSU 504.065, and from one parent at an additional 11 marker loci.

Description

BOTANICAL DENOMINATION

[0002] Corylus avellana

VARIETY DESIGNATION

[0003] `York`

BACKGROUND

[0004] The present Invention relates to a new and distinct cultivar of Corylus plant (hazelnut, filbert), botanically known as Corylus avellana, and hereinafter referred to by the name `York`. Corylus avellana is in the family Betulaceae.

[0005] The new Corylus resulted from a controlled cross of female parent OSU 479.027 (unpatented) and male parent OSU 504.065 (unpatented) made in 1997 by Shawn A. Mehlenbacher and David C. Smith. Hybrid seeds from the cross were harvested in August 1997, stratified, and seedlings grown in the greenhouse during the summer of 1998. From this cross, total of 183 seedling trees were planted in the field in Corvallis, Oreg., USA in October, 1998. `York` was discovered and selected by the Inventors as a single plant within the progeny of the stated cross-pollination in a controlled environment in Corvallis, Oreg.

[0006] `York` was originally assigned the designation OSU 878.048, which indicates the row and tree location of the original seedling. OSU 479.027 is from a cross of OSU 231.111 (unpatented)×OSU 226.122 (unpatented). OSU 504.065 is from a cross of OSU 186.080 (unpatented)×VR 17-15 (unpatented). The pedigree of `York` includes `Barcelona` (unpatented) widely grown in Oregon, `Casina` from Asturias, Spain (unpatented), `Tonda Gentile delle Langhe` from Piemonte, northern Italy (unpatented), `Montebello` from Sicily, Italy (unpatented), and `Tombul Ghiaghli`, a Turkish cultivar obtained from Greece (unpatented).

[0007] The new cultivar was asexually reproduced by rooted suckers annually for eight years (2003-2010) in Corvallis, Oreg. The new cultivar was also asexually propagated by whip grafting in Corvallis, Oreg. The unique features of this new Corylus are stable and reproduced true-to-type in successive generations of asexual reproduction.

SUMMARY

[0008] The following traits have been repeatedly observed and are determined to be the unique characteristics of `York`. These characteristics in combination distinguish `York` as a new and distinct cultivar:

[0009] 1. Globose plant habit and moderate vigor.

[0010] 2. Green developing and fully expanded leaves during the spring and summer.

[0011] 3. Resistance to eastern filbert blight caused by the fungus Anisogramma anomala (Peck) E. Muller.

[0012] 4. Presence of random amplified polymorphic DNA markers 152-800 and AAl12-850 in DNA of `York` amplified by the polymerase chain reaction. These two markers are linked to a dominant allele for resistance to eastern filbert blight from the cultivar Gasaway (unpatented).

[0013] 5. Expression of incompatibility alleles S₂ and S₂₁ in the styles.

[0014] 6. DNA fingerprints at 10 of 23 microsatellite marker loci differ from both parents OSU 479.027 and OSU 504.065, and from one parent at an additional 11 marker loci. The microsatellite primers are shown in Table 1, and allele sizes are shown in Table 2. DNA fingerprints of standard cultivars `Barcelona`, `Tonda Gentile delle Langhe` and `Extra Ghiaghli`, and `Gasaway`, the source of eastern filbert blight resistance, are also shown in attached Table 2.

[0015] In comparisons in two replicated trials conducted in Corvallis, Oreg., plants of the new Corylus differed from plants of the Corylus avellana cultivar Barcelona (unpatented), and other cultivars and selections of Corylus avellana known to the Inventors primarily in nut size, nut shape, kernel percentage (ratio of kernel weight to nut weight), frequency of blank nuts (nuts lacking kernels), time of pollen shed, time of nut maturity, length of the husk or involucre, and plant size.

BRIEF DESCRIPTION OF THE DRAWING

[0016] The accompanying colored photographs illustrate the overall appearance of the new cultivar, showing the colors as true as it is reasonably possible to obtain in colored reproductions of this type. Foliage colors in the photographs may differ slightly from the color values cited in the detailed botanical description which accurately describe the colors of the new Corylus.

[0017] FIG. 1 shows a tree of the new cultivar `York` growing in a field in August, in Corvallis, Oreg.

[0018] FIG. 2 shows the tree of the new cultivar `York` growing in a field in January, in Corvallis, Oreg.

[0019] FIG. 3 shows typical nuts, raw kernels, and blanched kernels of `York` hazelnut compared to those of `Jefferson` hazelnut.

[0020] FIG. 4 shows typical nuts, raw kernels, and blanched kernels of `York` hazelnut compared to those of `Lewis` hazelnut.

[0021] FIG. 5 shows the typical nuts, raw kernels, and blanched kernels of `York` hazelnut compared to those of `Barcelona` hazelnut and other hazelnut cultivars.

DETAILED BOTANICAL DESCRIPTION

[0022] The cultivar York has not been observed under all possible environmental conditions. The phenotype may vary somewhat with variations in environment such as temperature and light intensity, without, however, any variance in genotype. The aforementioned photographs and following observations and measurements describe plants grown in Corvallis, Oreg. under commercial practice outdoors in the field during the fall, winter and spring. Plants used for the photographs and description were propagated by tie-off layerage and growing on their own roots, and about seven years old. In the following description, color references are made to The Royal Horticultural Society Colour Chart, 1966 Edition, except where general terms of ordinary dictionary significance are used.

[0023] Botanical classification: Corylus avellana cultivar York.

[0024] Parentage:

[0025] Female, or seed, parent.--Corylus avellana selection OSU 479.027 (unpatented).

[0026] Male, or pollen, parent.--Corylus avellana selection OSU 504.065 (unpatented).

[0027] Propagation (type rooted suckers):

[0028] Time to initiate roots.--about 30 days at 20 degrees C.

[0029] Time to produce a rooted young plant.--about six months at 22 degrees C.

[0030] Root description.--fine to thick; freely branching; creamy white in color.

[0031] Propagation (type whip grafting):

[0032] Time to budbreak on the scions.--about 14 days at 25° C.

[0033] Time to produce a grafted plant.--about six months at 25° C.

[0034] Plant description:

[0035] General appearance.--perennial shrub.

[0036] Size.--plant height is about 5 meters; plant diameter or spread is about 5 meters.

[0037] Form.--spreading plant habit.

[0038] Growth and branching habit.--freely branching; about 15 lateral branches develop per plant. Pinching, i.e., removal of the terminal apices, enhances branching with lateral branches potentially forming at every node.

[0039] Vigor.--Moderate vigor growth habit.

[0040] Roots.--Fine to thick; freely branching; creamy white in color.

[0041] Lateral branch description:

[0042] Length.--about 38 cm.

[0043] Diameter.--about 6 mm.

[0044] Internode length.--about 3.2 cm.

[0045] Texture.--smooth, glabrous.

[0046] Strength.--strong.

[0047] Color.--immature -- 152B, mature -- 152B.

[0048] Foliage description:

[0049] Arrangement.--alternate, simple.

[0050] Length.--about 10.0 cm.

[0051] Width.--about 9.3 cm.

[0052] Shape.--ablong to ovate.

[0053] Apex.--obtuse to acute.

[0054] Base.--cordate.

[0055] Margin.--serrate.

[0056] Texture.--upper and lower surfaces -- slightly pubescent.

[0057] Venation pattern.--pinnate.

[0058] Color.--Developing foliage: upper surface 146B, lower surfaces: 146C. Fully expanded foliage, upper surface: Spring and summer, 146A; late summer and fall, 146A. Fully expanded foliage, lower surface: Spring and summer, 146C; late summer and fall, 146C. Venation, upper surface: Spring and summer, 146A; late summer and fall, 146A. Venation, lower surface: Spring and summer, 148D; late summer and fall, 148D.

[0059] Petiole description:

[0060] Length.--about 2.7 cm.

[0061] Diameter.--about 1.8 mm.

[0062] Texture.--upper and lower surfaces: pubescent.

[0063] Color.--upper surface: Spring and summer, 139D; late summer and fall, 139D. lower surface: Spring and summer, 139D; late summer and fall, 139D.

[0064] Flower description:

[0065] Male inflorescences.--catkins, color prior to elongation 194C.

[0066] Female inflorescence.--style color 047B.

[0067] Nut description:

[0068] Length.--about 18.0 mm.

[0069] Width.--about 19.7 mm.

[0070] Depth.--about 17.0 mm.

[0071] Nut shape.--round.

[0072] Nut shape index [(width+depth)/2*length].--1.02.

[0073] Nut compression index (width/depth).--1.16.

[0074] Nut shell color.--164A.

[0075] Nut weight.--about 2.73 grams.

[0076] Kernel weight.--about 1.23 grams.

[0077] Kernel percentage (kernel weight/nut weight).--about 46%.

[0078] Disease/pest resistance: Plants of the new Corylus are highly resistant to eastern filbert blight caused by the fungus Anisogramma anomala (Peck) E. Muller. Plants of the new Corylus are highly resistant to bud mites (Phytoptus avellanae Nal.), while plants of `Tonda Gentile delle Langhe` are highly susceptible, and plants of `Barcelona` are highly resistant.

[0079] Temperature tolerance: Tolerates temperatures from -10 to 38° C. in the field in Corvallis, Oreg.

TABLE-US-00001

[0079] TABLE 1 Primers and annealing temperatures for the 24 microsatellite marker loci used to fingerprint `York` and other hazelnut cultivars. Repeat Locus motif Size T_n n He Ho PIC r A613 (TC)₁₃ 149- 60 14 0.85 0.85 0.85 0.00 (CA)₁₂ 177 A614 (TC)₁₇ 125- 60 14 0.85 0.85 0.84 0.00 (CA)₁₀ 156 NNN (CA)₆ A616 (AC)₁₁ 136- 60 13 0.85 0.85 0.83 0.00 162 A640 (CT)₁₅ 354- 67 11 0.80 0.73 0.7 0.04 (CA)₁₃ 378 B107 (CT)₁₄ 112- 55 14 0.85 0.80 0.83 0.02 151 B617 (GA)₁₅ 280- 60 9 0.80 0.78 0.78 0.01 298 B619 (TC)₂₁ 146- 60 14 0.88 0.88 0.7 0.00 180 B634 (AG)₁₅ 218- 60 9 0.76 0.76 0.73 0.00 238 B657 (AG)₁₅ 210- 60 8 0.84 0.98 0.82 -0.08 228 B671 (AG) 221- 60 13 0.86 0.88 0.84 -0.01 6NN 249 (GA)₁₇ B709 (GA)₂₁ 219- 60 8 0.74 0.76 0.70 -0.01 233 B733 (TC)₁₅ 161- 60 8 0.68 0.68 0.63 0.00 183 B741 (GT)₅ 176- 60 10 0.77 0.78 0.74 0.00 (GA)₁₂ 194 B749 (TC)₁₂ 200- 60 6 0.60 0.64 0.51 -0.03 210 B751 (GA)₁₅ 141- 60 7 0.80 0.80 0.77 0.01 153 B774 (AG)₁₅ 195- 60 8 0.80 0.80 0.77 0.01 213 B776 (GA)₁₇ 134- 60 7 0.71 0.60 0.67 0.07 148 B795 (TC)₈ 296- 60 12 0.76 0.74 0.74 0.01 Ns 332 (CT)₇ Ns(CT) ₁₀Ns (TC)₅ C115 (TAA)₅ 167- 60 14 0.80 0.80 0.77 0.00 (GAA)₁₂ 226 KG809 (AGG)₆ 333- 55 5 0.66 0.64 0.60 0.01 345 KG811 (GA)₁₇ 240- 58 12 0.83 0.82 0.81 0.01 278 KG827 (CT)₁₃ 264- 67 9 0.78 0.84 0.75 -0.04 AA 282 (CA)₇ KG830 (CT)₁₄ 279- 67 9 0.79 0.78 0.76 0.00 GTATT 311 (CA)₈ Soman- (AAT)₅ 54 3 0.60 0.98 0.51 -0.27 G Primers Primers Refer- Locus LG 5'-3' 5'-3' ence A613 11 Ned- R- Gurcan CACACG CCCCTT et al. CCTTGTCA TCACATGT 2010 CTCTTT TTGCTT (SEQ ID (SEQ ID NO: 1) NO: 2) A614 6 Hex- R- Gurcan TGGCAGA GCAGTGG et al. GCTTTGT AGGATTG 2010 CAGCTT CTGACT (SEQ ID (SEQ ID NO: 3) NO: 4) A616 8 Fam- R- Gurcan CACTCAT ATGGCTT et al. ACCGCAA TTGCTTC 2010 ACTCCA GTTTTG (SEQ ID (SEQ ID NO: 5) NO: 6) A640 10 F- Fam- Gurcan TGCCTCT CGCCATAT et al. GCAGTTA AATTGGGAT 2010 GTCATCAA GCTTGTTG ATGTAGG (SEQ ID (SEQ ID NO: 8) NO: 7) B107 10 Ned- R- Boccacci GTAGGTGC AACACCATA et al. ACTTGATG TTGAGTCT 2005; TGCTTTAC TTCAAAGC Go- (SEQ ID (SEQ ID kirmak NO: 9) NO: 10) et al. 2009 B617 8 Fam- R- Gurcan TCCGTGT TGTTTT et al. TGAGTAT TGGTGGA 2010 GGACGA GCGATG (SEQ ID (SEQ ID NO: 11) NO: 12) B619 3 Fam- R- Gurcan AGTCGG GCGATC et al. CTCCCCT TGACCTC 2010 TTTCTC ATTTTTG (SEQ ID (SEQ ID NO: 13) NO: 14) B634 4 Hex- R- Gurcan CCTGCA GTGCAG et al. TCCAGGA AGGTTGC 2010 CTCATTA ACTCAAA (SEQ ID (SEQ ID NO: 15) NO: 16) B657 11 Ned- R- Gurcan GAGAGT AGCCTC et al. GCGTCTT CAACCTC 2010 CCTCTGG ACGAAC (SEQ ID (SEQ ID NO: 17) NO: 18) B671 9 Hex- R- Gurcan TTGCCAG ACCAGC et al. TGCATAC TCTGGGC 2010 TCTGATG TTAACAC (SEQ ID (SEQ ID NO: 19) NO: 20) B709 5 Ned- R- Gurcan CCAAGCA GCGGGT et al. CGAATGA TCTCGTT 2010 ACTCAA GTACACT (SEQ ID (SEQ ID NO: 21) NO: 22) B733 7.2 Ned- R- Gurcan CACCCT CATCCC et al. CTTCACC CTGTTGG 2010 ACCTCAT AGTTTTC (SEQ ID (SEQ ID NO: 23) NO: 24) B741 5 Fam- R- Gurcan GTTCACA CGTGTT et al. GGCTGTT GCTCATG 2010 GGGTTT TGTTGTG (SEQ ID (SEQ ID NO: 25) NO: 26) B749 1 Hex- R- Gurcan GGCTGA TCGGCTA et al. CAACACA GGGTTAG 2010 GCAGAAA GGTTTT (SEQ ID (SEQ ID NO: 27) NO: 28 B751 7.2 Fam- R- Gurcan AGCTGGT AAACTCAA et al. TCTTCGA ATAAAACC 2010 CATTCC CCTGCTC (SEQ ID (SEQ ID NO: 29) NO: 30) B774 5 Ned- R- Gurcan GTTTTG TGTGTGT et al. CGAGCTC GGTCTGT 2010 ATTGTCA AGGCACT (SEQ ID (SEQ ID NO: 31) NO: 32) B776 6 Fam- R- Gurcan TGTATGTA TGAGGGG et al. CACACGGA AAGAGGT 2010 GAGAGAGA TTGATG (SEQ ID (SEQ ID NO: 33) NO: 34) B795 NA Fam- R- Gurcan GACCCACA TGGGCA et al. AACAATAA TCATCCA 2010 CCTATCTC GGTCTA (SEQ ID (SEQ ID NO: 35) NO: 36) C115 4 Fam- GTTTCCAG Bassil A1111 CC ATCTGCCTC et al. GCAGATA CATATAAT 2005b, ATACAGG (SEQ ID Go- (SEQ ID NO: 38) kirmak NO: 37) et al. 2009 KG809 4 Hex- F- Gurcan AGGCAT GGAAGGT and CAGTTC GAGAGAA Mehlen- ATCCAA ATCAAGT bacher (SEQ ID (SEQ ID 2010 NO: 39) NO: 40) KG811 2 Ned- F- Gurcan AAGGCG GAACAAC and GCACTC TGAAGAC Mehlen- GCTCACA GCAAAG bacher (SEQ ID (SEQ ID 2010 NO: 41) NO: 42) KG827 9 Fam- GAGGGA Gurcan AGAACTCC GCAAGTCA and GACTAATA AAGTTGA Mehlen- ATCCTAA GAAGAAA bacher CCCTTGC (SEQ ID 2010 (SEQ ID NO: 44) NO: 43)

KG830 9 Ned- AAAGCA Gurcan TGGAGGA ACTCA and AGTTTTGA TAGCT Mehlen- ATGGTAG GAAGTC bacher TAGAGGA CAATCA 2010 (SEQ ID (SEQ ID NO: 45) NO: 46) Soman-G NA Hex- R- unpub- TGGCGT GCCATCTTT lished TGCAACA AGAAAGTTC TATTCTC GATACAG (SEQ ID (SEQ ID NO: 47) NO: 48) Primer fluorescent tags are FAM, HEX, and NED. Ta: annealing-temperature (° C.) N: number of alleles He: expected heterozygosity Ho: observed heterozygosity PIC: polymorphism information content r: estimated null allele frequency LG: linkage group

TABLE-US-00002 TABLE 2 Allele sizes in `York`, its parents, and four other hazelnut cultivars at 24 microsatellite loci. `Tonda Gentile Tag Locus `York` 479.027 504.065 delle Langhe` NED A613 157/177 177/177 157/159 151/157 HEX A614 125/158 125/132 148/158 125/135 FAM A616 142/150 142/150 150/150 148/150 FAM A640 362/374 354/374 354/362 354/368 NED B107 122/134 134/146 122/134 134/152 FAM B617 286/290 286/290 286/286 286/296 FAM B619 156/164 156/164 156/164 148/164 HEX B634 226/234 226/226 232/234 226/226 NED B657 220/222 222/226 220/224 218/226 HEX B671 241/247 227/247 241/241 237/241 NED B709 227/231 227/231 227/227 227/227 NED B733 171/179 171/179 173/179 171/173 FAM B741 177/186 177/186 177/184 177/184 HEX B749 208/208 206/208 206/208 206/208 FAM B751 151/153 151/153 143/151 149/153 NED B774 203/209 209/211 203/207 203/211 FAM B776 137/150 137/137 137/150 137/137 FAM B795 330/330 296/330 330/330 312/330 FAM C115 197/197 194/197 194/197 173/173 HEX KG809 336/345 336/339 339/345 336/339 NED KG811 254/254 254/254 254/264 254/264 FAM KG827 266/270 270/282 266/266 266/268 NED KG830 295/295 295/295 295/295 291/295 HEX SMNG 196/200 196/200 196/200 196/200 `Extra Tag `Barcelona` Ghiaghli` `Gasaway` NED 151/159 167/169 159/161 HEX 125/131 125/150 143/158 FAM 142/150 150/158 148/148 FAM 354/374 374/374 362/368 NED 112/134 116/116 122/128 FAM 286/290 294/296 292/296 FAM 156/170 164/174 170/174 HEX 226/226 226/226 220/232 NED 218/222 210/222 224/228 HEX 223/227 227/247 235/247 NED 225/233 225/227 227/227 NED 171/173 171/171 173/173 FAM 177/186 177/184 186/188 HEX 208/208 208/208 206/208 FAM 143/153 143/147 143/143 NED 203/207 195/203 203/209 FAM 135/137 135/137 146/150 FAM 330/330 296/310 314/316 FAM 173/194 182/194 215/218 HEX 336/336 336/339 336/345 NED 258/264 240/242 254/258 FAM 280/282 276/282 270/280 NED 291/295 291/295 291/305 HEX 196/200 196/200 196/196

REFERENCES

[0080] Bassil N. V., Botta R., Mehlenbacher S. A. 2005a. Microsatellite markers in hazelnut: Isolation, characterization and cross-species amplification. J. Amer. Soc. Hort. Sci. 130:543-549.

[0081] Bassil N. V., Botta R., Mehlenbacher S. A. 2005b. Additional microsatellite markers of the European hazelnut. Acta Hort. 686:105-110.

[0082] Boccacci P., Akkak A., Bassil N. V., Mehlenbacher S. A., Botta R. 2005. Characterization and evaluation of microsatellite loci in European hazelnut (C. avellana) and their transferability to other Corylus species. Molec. Ecol. Notes 5:934-937.

[0083] Boccacci P., Akkak, A. and Botta, R. 2006. DNA typing and genetic relations among European hazelnut (Corylus avellana L.) cultivars using microsatellite markers. Genome 49:598-611.

[0084] Gokirmak T., Mehlenbacher S. A., Bassil N. V. 2009. Characterization of European hazelnut (Corylus avellana) cultivars using SSR markers. Genetic Resources and Crop Evolution 56:147-172.

[0085] Gurcan, K., S. A. Mehlenbacher and V. Erdogan. 2010a. Genetic diversity in hazelnut cultivars from Black Sea countries assessed using SSR markers. Plant Breeding (available on-line doi:10.1111/j.1439-0523.2009.01753.x).

[0086] Gurcan, K., S. A. Mehlenbacher, N. V. Bassil, P. Boccacci, A. Akkak and R. Botta. 2010b. New microsatellite markers for Corylus avellana from enriched libraries. Tree Genetics and Genomes (available on-line as DOI 10.1007/s11295-010-0269-y).

[0087] Gurcan, K. and S. A. Mehlenbacher. 2010. Development of microsatellite marker loci for European hazelnut (Corylus avellana L.) from ISSR fragments. Molecular

[0088] Breeding (available on-line).

Sequence CWU 1

1

48120DNAArtificial SequenceSynthetic polynucleotide 1cacacgcctt gtcactcttt 20220DNAArtificial SequenceSynthetic polynucleotide 2cccctttcac atgtttgctt 20320DNAArtificial SequenceSynthetic polynucleotide 3tggcagagct ttgtcagctt 20420DNAArtificial SequenceSynthetic polynucleotide 4gcagtggagg attgctgact 20520DNAArtificial SequenceSynthetic polynucleotide 5cactcatacc gcaaactcca 20620DNAArtificial SequenceSynthetic polynucleotide 6atggcttttg cttcgttttg 20729DNAArtificial SequenceSynthetic polynucleotide 7tgcctctgca gttagtcatc aaatgtagg 29825DNAArtificial SequenceSynthetic polynucleotide 8cgccatataa ttgggatgct tgttg 25924DNAArtificial SequenceSynthetic polynucleotide 9gtaggtgcac ttgatgtgct ttac 241025DNAArtificial SequenceSynthetic polynucleotide 10aacaccatat tgagtctttc aaagc 251120DNAArtificial SequenceSynthetic polynucleotide 11tccgtgttga gtatggacga 201219DNAArtificial SequenceSynthetic polynucleotide 12tgtttttggt ggagcgatg 191319DNAArtificial SequenceSynthetic polynucleotide 13agtcggctcc ccttttctc 191420DNAArtificial SequenceSynthetic polynucleotide 14gcgatctgac ctcatttttg 201520DNAArtificial SequenceSynthetic polynucleotide 15cctgcatcca ggactcatta 201620DNAArtificial SequenceSynthetic polynucleotide 16gtgcagaggt tgcactcaaa 201720DNAArtificial SequenceSynthetic polynucleotide 17gagagtgcgt cttcctctgg 201819DNAArtificial SequenceSynthetic polynucleotide 18agcctcacct ccaacgaac 191921DNAArtificial SequenceSynthetic polynucleotide 19ttgccagtgc atactctgat g 212020DNAArtificial SequenceSynthetic polynucleotide 20accagctctg ggcttaacac 202120DNAArtificial SequenceSynthetic polynucleotide 21ccaagcacga atgaactcaa 202220DNAArtificial SequenceSynthetic polynucleotide 22gcgggttctc gttgtacact 202320DNAArtificial SequenceSynthetic polynucleotide 23caccctcttc accacctcat 202420DNAArtificial SequenceSynthetic polynucleotide 24catcccctgt tggagttttc 202520DNAArtificial SequenceSynthetic polynucleotide 25gttcacaggc tgttgggttt 202620DNAArtificial SequenceSynthetic polynucleotide 26cgtgttgctc atgtgttgtg 202720DNAArtificial SequenceSynthetic polynucleotide 27ggctgacaac acagcagaaa 202820DNAArtificial SequenceSynthetic polynucleotide 28tcggctaggg ttagggtttt 202920DNAArtificial SequenceSynthetic polynucleotide 29agctggttct tcgacattcc 203023DNAArtificial SequenceSynthetic polynucleotide 30aaactcaaat aaaacccctg ctc 233120DNAArtificial SequenceSynthetic polynucleotide 31gttttgcgag ctcattgtca 203221DNAArtificial SequenceSynthetic polynucleotide 32tgtgtgtggt ctgtaggcac t 213324DNAArtificial SequenceSynthetic polynucleotide 33tgtatgtaca cacggagaga gaga 243420DNAArtificial SequenceSynthetic polynucleotide 34tgaggggaag aggtttgatg 203524DNAArtificial SequenceSynthetic polynucleotide 35gacccacaaa caataaccta tctc 243619DNAArtificial SequenceSynthetic polynucleotide 36tgggcatcat ccaggtcta 193721DNAArtificial SequenceSynthetic polynucleotide 37attttccgca gataatacag g 213825DNAArtificial SequenceSynthetic polynucleotide 38gtttccagat ctgcctccat ataat 253918DNAArtificial SequenceSynthetic polynucleotide 39aggcatcagt tcatccaa 184021DNAArtificial SequenceSynthetic polynucleotide 40ggaaggtgag agaaatcaag t 214118DNAArtificial SequenceSynthetic polynucleotide 41aaggcggcac tcgctcac 184221DNAArtificial SequenceSynthetic polynucleotide 42gaacaactga agacagcaaa g 214330DNAArtificial SequenceSynthetic polynucleotide 43agaactccga ctaataatcc taacccttgc 304428DNAArtificial SequenceSynthetic polynucleotide 44gagggagcaa gtcaaagttg agaagaaa 284529DNAArtificial SequenceSynthetic polynucleotide 45tggaggaagt tttgaatggt agtagagga 294628DNAArtificial SequenceSynthetic polynucleotide 46aaagcaactc atagctgaag tccaatca 284720DNAArtificial SequenceSynthetic polynucleotide 47tggcgttgca acatattctc 204825DNAArtificial SequenceSynthetic polynucleotide 48gccatcttta gaaagttcga tacag 25

Claims

1. A new and distinct cultivar of Corylus plant named `York`, as illustrated and described.

Images